Complete Chromatic Nevron Hunting Guide: Locations, Strategies & Rewards in Clair Obscur: Expedition 33

Chromatic Nevrons are the ultimate challenge in Clair Obscur: Expedition 33, offering the game's best loot for those brave enough to hunt them. These super-charged, rainbow-tinted variants of regular foes are scattered across the Continent, demanding mastery of combat and exploration. This guide provides the exact locations, strategies, and rewards you need to conquer every one and claim their elite gear.

What Are Chromatic Nevrons and Why Hunt Them?

Chromatic Nevrons: super-charged, rainbow-tinted variants of regular foes that roam Clair Obscur: Expedition 33. Each one is a late-game powerhouse, sporting bloated health bars and punishing move-sets that can flatten an unprepared party in seconds.

Guaranteed High-Tier Loot

Every Chromatic Nevron you defeat drops a fixed cache of top-shelf rewards:

• Elite Weapons – examples include Lune's Angerim (fire-based) and Sciel's Moisson (dark damage conversion), both among the strongest arms in the game.
• Rare Pictos – defensive gems like Anti-Stun and Anti-Blight that grant full immunity to their respective status effects.
• Upgrade Materials & Tints – scarce resources needed to push gear to its final tier.

Completionist Necessity

Nearly 50 of these optional bosses are scattered across the Continent, many tucked into hidden corners of zones like Renoir's Drafts or the Flying Manor. Tracking them all demands thorough exploration - some require flight, others underwater traversal - but the payoff is twofold: access to the game's best gear and the trophies tied to 100 % completion.

Essential Hunting Strategies and Preparation

Chromatic Nevron Hunting Tips

Chromatic Nevrons are among the most formidable optional bosses in Clair Obscur: Expedition 33, offering some of the best loot in exchange for significant challenge. With nearly 50 different enemies scattered across the Continent, systematic searching is essential.

• Start Small: Begin with Chromatic Bruler, noted as the easiest target and found early in the game to build confidence and gain rewards
• Level Appropriately: Early Chromatic Nevrons may be excessively challenging for under-leveled parties - improve gear before attempting difficult encounters
• Explore Thoroughly: Many are hidden in less-explored areas easily overlooked by players, requiring careful examination of every corner
• Master Mobility: Later encounters require specific skills - some need flight abilities, while Chromatic Creation demands Esquie's underwater diving capabilities
• Test Elements: Exploit weaknesses by experimenting with damage types, though the game doesn't always follow predictable rules (fire-based Chromatic Luster is weak to Lightning)
• Use Community Knowledge: Consult location guides for systematic hunting across all zones

Expedition 33 Rare Enemy Strategy

Rare enemy hunting in Expedition 33 requires a methodical approach combining strategic exploration with optimal party composition. Success depends more on preparation than brute force.

Fast Travel and Respawn Mechanics

Expedition Flags serve as your primary fast travel system, acting as reference points for exploration. You can only fast-travel from these designated rest points, making their placement crucial for efficient hunting. When accessing an Expedition Flag, all previously defeated enemies in the area will respawn, creating opportunities for repeated encounters.

Party Optimization

Focus on gear upgrades and skill unlocks rather than traditional level grinding. Lune stands out as the best sub-DPS for rare enemy hunting, offering Pyre, Frost, and Volt damage while providing healing, debuffs, and revival capabilities.

Systematic Exploration Process

• Backtrack with New Abilities: Many hidden areas containing rare enemies are gated behind skills unlocked later in the game
• Exhaust NPC Dialogue: Characters often provide cryptic hints about nearby secrets and hidden entrances
• Trigger Side Quests: These frequently lead to rare enemy areas not accessible through standard exploration
• Exploit Weaknesses: Almost every enemy has at least one weakness and one resistance - elemental damage dramatically increases effectiveness

Continent Overworld Chromatic Nevrons (Early-Mid Game)

These optional bosses roam The Continent's open world and drop powerful Pictos or unique weapons. All require Act 2+ progression and Esquie's flight or swim to reach the floating or offshore arenas.

Chromatic Lancelier – Spring Meadows

Location	Reward	Strategy
Grand Meadow rest point → right past rope → rock formation	Augmented Attack Picto (+50 % Base Attack as Lumina, +Defense/Speed) plus Gustave's Lanceram +1, 2 Chroma Catalysts, 540 Chroma	Strike the weak point immediately to avoid its long lunge combo.

Chromatic Goblu – North of Sky Island

Location	Reward	Strategy
Floating island NE of Coastal Cave; fly with Esquie	Gobluson (unique weapon for Sciel)	Exploit Fire and Poison weaknesses; keep distance from its spinning slam.

Chromatic Petank – East of Stone Wave Cliffs Cave

Location	Reward	Strategy
Valley NE of Stone Wave Cliffs Cave; lure to stone arch	Anti-Stun Picto, 2 Grandiose Chroma Catalysts, 5 Colour of Lumina	No weak points - clear summoned Nevrons first, then burst the boss.

Chromatic Aberration – South of the Fountain

Location	Reward	Strategy
Floating island SE of The Fountain; fly required	Longer Burn Picto (+2 Burn turns) + Brumaro for Monoco	Lvl-80 superboss; manage Burn stacks and dodge wide laser sweeps.

Chromatic Demineur – North of Stone Quarry

Location	Reward	Strategy
Island N of Stone Quarry; fly required	Angerim (Fire weapon for Lune, scales Luck/Defense)	Floating enemy—use elemental attacks and sidestep fire waves.

Chromatic Benisseur – East of Stone Wave Cliffs

Location	Reward	Strategy
Far-east mountains; pour Chroma into fountain	Benisim (Lvl-28 weapon for Lune)	Weak to Ice & Dark; parry spear thrusts for counter windows.

Chromatic Portier – Southeast Floating Island

Location	Reward	Strategy
Small island SE of Stone Wave Cliffs; fly required	Anti-Blight Picto (Blight immunity, +Health/Defense) + 2 Grandiose Catalysts	Three-phase fight—pack Blight cleanse or Verso's Burden.

Chromatic Jar – West of Gestral Village Island

Location	Reward	Strategy
SW island off Gestral Village; swim with Esquie	Jarum (weapon drop)	Resists physical; lean on elemental spells and status effects.

Chromatic Bruler – North of Gestral Village

Location	Reward	Strategy
Fields N of Gestral Village; circle red grass patches	Powerful Revive Picto (Powerful buff on revive, +Speed/Crit) + 21 k XP, 3 k Chroma, 2 Polished Catalysts	Act-1 viable; interrupt flame pillars and burst during overheat.

Chromatic Reaper Cultist – Northeast Coastal Cave Island

Location	Reward	Strategy
Island NE of Coastal Cave; swim required	Blodam (unique weapon for Sciel)	Flying foe weak to Dark; later re-fightable on Endless Tower 10-1.

Chromatic Moissonneuse – Southwest Visages Island

Location	Reward	Strategy
Island SW of Visages, near Old Lumière	Moisson (Agility/Luck scythe for Sciel, Twilight trait: Dark damage +20 %)	Apply Burn, break its bar, then unload Fire/Dark burst during stun.

Dungeon and Story Area Chromatic Nevrons

Chromatic Danseuse – Old Lumiere

The Chromatic Danseuse awaits in the Train Station Ruins of Old Lumiere, but only after you have bested Renoir in Act 2. Expect a punishing fight—level 33 or higher is strongly advised.

Location	Reward	Key Strategy
Old Lumiere Train Station Ruins	Revive Paradox (ability)	Exploit Ice weakness; bring ice-focused characters and abilities. The boss carries a "ridiculous" health pool, so plan for a long encounter.

Chromatic Abbest – Abbest Cave

Early explorers can tackle Chromatic Abbest as soon as they unlock world-map travel. Found inside Abbest Cave, this encounter is designed to test your grasp of combat fundamentals.

Location	Reward	Key Strategy
Abbest Cave, first region	First Offensive (ability)	Target Wind/Lightning and Slash damage; avoid Strike or Pierce. Watch for Chromatic Barrage, Petal Storm, and Charge & Slam patterns.

Chromatic Hexga – Stone Wave Cliffs Cave

Swim east from Stone Wave Cliffs after Act 2 to reach the hidden cave housing Chromatic Hexga. Its shoulder crystal changes color to telegraph escalating danger.

Location	Reward	Key Strategy
Stone Wave Cliffs Cave	Energising Revive OR Confident (choose one)	Resists Fire; burst it down before the red-crystal phase. Victory yields 3 Pictos for upgrades.

Chromatic Chalier – Sinister Cave

Far east of Sirene, on a tiny island near Gestral Beach, Chromatic Chalier guards the Sinister Cave. Consistent Burn application is the key to victory.

Location	Reward	Key Strategy
Sinister Cave	Chalium (Maelle's unique weapon)	Weak to Ice & Fire; maintain Burn status. Learn to dodge Swiping, Relentless, and Punching combos.

Chromatic Chapelier – The Crows

A secret encounter, Chromatic Chapelier spawns only after you shoot all 25 glowing crows in The Crows area. Mastery of the parry mechanic is mandatory.

Location	Reward	Key Strategy
The Crows, northwest Continent	Shielding Death (ability)	Parry the opening mask volley perfectly; the entire fight revolves around timed counters.

Chromatic Luster – Forgotten Battlefield

Drop down to the right of the Fading Woman from the Battlefield checkpoint to face Chromatic Luster. Fire heals it—leave your pyromaniacs at home.

Location	Reward	Key Strategy
Forgotten Battlefield underground	Lusteson (Sciel weapon) + Energising Parry	Exploit Lightning weakness; avoid Fire at all costs. Expect a three-hit Fast Combo that inflicts Burn.

Chromatic Cruler – Endless Night Sanctuary

Accessible only after Act 3's flight unlock, Chromatic Cruler waits beyond the Night Totem flag. Its limited move set is deceptively simple.

Location	Reward	Key Strategy
Endless Night Sanctuary, Night Totem area	Cruleram weapon + Quick Break Pictos	Resists Lightning; pack Burn/Stun cures. A lamp-pole melee pattern leaves clear dodge windows.

Chromatic Bourgeon – The Monolith

Hidden left of the Tainted Waters flag behind seaweed on a hillside, Chromatic Bourgeon is considered the ultimate pre-Paintress test.

Location	Reward	Key Strategy
The Monolith, Tainted Waters	Bourgelon (top-tier Sciel weapon)	Master dodge-and-counter timing; varied, dangerous combos demand flawless execution.

Chromatic Braseleur – The Reacher

Grapple up from The Peak flagstone to Alicia's summit for Chromatic Braseleur, a fight tied to Maelle's personal quest.

Location	Reward	Key Strategy
The Reacher peak	Braselim (unique weapon)	Weak to Fire & Lightning, resists Frost/Physical. Watch for Stance Switch and the devastating 6-hit Hammer Combo.

Chromatic Gault – Stone Wave Cliffs

Behind a Paint Spike in the Old Farm area, Chromatic Gault requires the Paint Break ability earned by collecting four of Sastro's Lost Gestrals.

Location	Reward	Key Strategy
Stone Wave Cliffs, Old Farm behind Paint Spike	Tainted (weapon)	Sciel excels here; frequent AoE party-wide hits demand strong healing and tanking. Farm Colour of Lumina afterward.

Chromatic Ramasseur – Visages: Sadness Vale

Just off the main path in Visages lies Sadness Vale, where Chromatic Ramasseur forces a "parry-only" duel.

Location	Reward	Key Strategy
Visages, Sadness Vale	Augmented Counter II	Perfect parries create openings; exploit Light weakness with Verso's Perfect Break and Purification. Rewards include 4 410 Chroma, 61 598 XP, and 5× Colour of Lumina.

Special and Endgame Area Chromatic Nevrons

Below is a quick-reference atlas to every Chromatic Nevron tucked away in Clair Obscur's toughest corners. Each entry lists where to find the fight, the exact loot you'll earn, and the single most important tactic to survive.

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Chromatic Clair Obscur – Endless Tower Stage 11

Location	Reward	Strategy
Endless Tower Stage 11, Trial 2 – summit of The Monolith	1× Grandiose Chroma Catalyst, 1× Colour of Lumina	Defeat Lampmaster and Creation first to soften the final encounter, then exploit its rotating elemental stance by matching the current weakness.

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Chromatic Creation – Renoir's Drafts

Location	Reward	Strategy
Act 3 sea-zone "Renoir's Drafts"; grapple up the stairs	1× Perfect Chroma Catalyst, 2× Grandiose Chroma Catalyst, 5× Colour of Lumina	Prepare for a massive difficulty spike—this is one of the deadliest fights in the game.

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Chromatic Boucheclier – Isle of Eyes

Location	Reward	Strategy
North of Visages; solve the eye-shooting puzzle to restore the bridge	Facesum (Maelle, Lv 22), 2× Grandiose Chroma Catalyst, 1× Colour of Prism	No weak spot—learn to dodge Prismatic Barrage, Spectrum Slam, and Chromatic Wave.

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Chromatic Glaise – Sky Island

Location	Reward	Strategy
East of Coastal Cave, reachable with Esquie's flight; also Endless Tower Stage 8-3	Urnaro (Monoco, Lv 20 Earth weapon), 2× Grandiose Chroma Catalyst, 5× Colour of Lumina	Vitality- and Luck-scaling weapon that speeds up Almighty Mask access.

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Chromatic Glissando – Sirène's Dress

Location	Reward	Strategy
Sirène's Dress (requires flight); part of Lune's relationship quest	Choralim (Lv 4 weapon)	Two-phase fight: worm first, then three Dark Ballet adds.

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Chromatic Gold Chevalière – Crimson Forest

Location	Reward	Strategy
Southwest of Blades' Graveyard; activate three statues to spawn the boss	Perilous Parry Picto	Bring both Light and Dark damage—Clair (weak to Dark) and Obscur (weak to Light) assist the boss.

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Chromatic Greatsword Cultist – Sirène

Location	Reward	Strategy
Inside the Sirène dungeon during the main quest	Sireso (greatsword for Gustave/Verso), 2× Resplendent Chroma Catalyst, 5× Colour of Lumina	Stack Dark-element attacks; it resists Light.

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Chromatic Orphelin – Yellow Harvest

Location	Reward	Strategy
End of Act 1 optional zone Yellow Harvest	Kralim (Lune, Lv 7 Lightning weapon), 6× Polished Chroma Catalyst, 15× Colour of Lumina, 54 429 XP, 7 988 Chroma	Trio of Nevrons wielding kitchen utensils—watch the stacking Blight.

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Chromatic Potier – Floating Island (Act 3)

Location	Reward	Strategy
Overworld floating island in Act 3	Potierim	Weak to Sword and Wind; break its legs to bypass damage reduction. Also appears in Endless Tower 7-2 & 10-2.

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Chromatic Sapling – Falling Leaves

Location	Reward	Strategy
Western edge of the Continent in Act 2	Saperim (Lune, Lv 4 Lightning weapon)	Absorbs Dark—use any other element.

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Chromatic Stalact – Frozen Hearts

Location	Reward	Strategy
Frozen Hearts optional zone	Stalum (Lv 21 weapon upgrade)	Farm regular Stalact groups for up to 3 upgrade drops if you miss the boss.

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Chromatic Veilleur – Frozen Hearts

Location	Reward	Strategy
Large cavern near Glacial Falls; enter via ice tunnel after picking up Burn Affinity	Energising Burn Picto (+1 AP when inflicting Burn, once per turn), 2× Grandiose Chroma Catalyst, 5× Colour of Lumina, 10 448 Chroma	Survive the opening Blighted Wave (6 stacks of Blight) and exploit Burn/Stun weaknesses.

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Chromatic Troubadour – Flying Waters

Location	Reward	Strategy
Upper path of Flying Waters; climb left-hand holds from Lumièran Streets	Troubadim (Lv 4 weapon), 2× Chroma Catalyst, 5× Colour of Lumina	Weak to Light; aim for the small dot on its chest. Watch the deceptive Tricky Shot timing.

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Chromatic Échassier – Lumière

Location	Reward	Strategy
Main story path in Lumière during Act 3	Augmented Counter III (Offensive Picto: +75 % counter damage), 2× Resplendent Chroma Catalyst, 5× Colour of Lumina	Fire-element weakness; break its legs to remove damage reduction.

Arena and Multi-Boss Chromatic Encounters

Dark Gestral Arena Trio

The Dark Gestral Arena sits on a floating island and serves as an optional boss gauntlet. You pick one character to battle three Chromatic Sakapatate variants in sequence.

Enemy	Weakness	Reward
Chromatic Catapult Sakapatate	Fire	Level 25 Trebuchim (Lune)
Chromatic Ranger Sakapatate	Fire	—
Chromatic Robust Sakapatate	Wind	Level 25 Sakaram (Gustave/Verso)

Note: Ranger Sakapatate also resists Lightning damage.

Chromatic Ceramic Chevaliere - Duel Encounter

Found in Act 2's Old Lumière region, the Chromatic Ceramic Chevalière is an optional duel boss. It can appear alongside Steel Chevalieres, most notably on Endless Tower Stage 10.

Detail	Info
Location	Old Lumière (Act 2)
Weakness	Physical
Key Attack	Gradient Strike (second hit deals Gradient damage)
Recommended Skill	Phantom Strike (Maelle's Void AoE, Act III)

Standard Ceramic and Steel Chevalieres drop the Battlum weapon.

Chromatic Dualliste - Supercharged Variant

Marked by a name bar, the Chromatic Dualliste is a supercharged optional boss with a unique resistance profile.

Detail	Info
Resistances	Fire, Dark
Weakness	Light
Neutral to	Physical, Ice, Lightning, Earth, Void
Reward	Monoco skill Duallist Storm

The regular Dualliste in the Forgotten Battlefield is also weak to Light, making Verso invaluable.

Missing and Unconfirmed Chromatic Nevrons

Chromatic Ceramic Chevaliere - Location Pending

Confirmed to exist, but its exact location is still being verified. Current status: Deltia's Gaming marks the entry with "We will update this section once we find the Chromatic Nevron."
GameFAQs, however, places the fight in Endless Tower Stage 10-2, so the discrepancy is under review.

Chromatic Dualliste - Location Pending

The Chromatic variant is acknowledged, yet no source pinpoints where to trigger it. Standard Dualliste appears as a mandatory Act 2 boss at the end of the Forgotten Battlefield, but the Chromatic version remains undocumented.

Chromatic Eveque - Unverified

Listed by the Expedition 33 Wiki as an optional boss, Chromatic Eveque is believed to inhabit Spring Meadows—the same zone as the first major story fight. Until a first-hand sighting surfaces, treat this as unconfirmed.

Chromatic Gargant - Location Known, Reward: Garganon

Location	Reward
Island southwest of Visages Continent, Moisson region	Garganon

Note: some guides confuse this spot with Chromatic Moissonneuse, who actually occupies the west side of the same island and drops 3,885 Chroma.

Chromatic Mage Cultist - Location Pending

Weaknesses are already catalogued—Dark and Light damage—but the arena itself is missing. Deltia's entry reads "We will update this section once we find the Chromatic Nevron."
Confirmed drop: 1× Duenum.

Chromatic Pelerin - Location Pending

Only placeholder text exists. No community reports or datamined coordinates have surfaced yet.

Chromatic Steel Chevaliere - No Weapon Drop

Found in Endless Tower Stage 10-2 alongside Chromatic Ceramic and Gold Chevalieres, this triple-boss fight is unique for not yielding any weapon or Pictos. Its only notable trait is a Physical weakness.

Chromatic Volester - Unverified

No reliable source has documented this variant. Standard Volesters first appear in Act I Spring Meadows' Grand Meadow and are weak to Ice, yet the Chromatic form remains elusive.

Reward Analysis: Best Weapons and Pictos from Chromatic Nevrons

Best Weapons Tier List

Chromatic Nevrons are elite enemies that drop many of the best weapons in Clair Obscur: Expedition 33. These drops can dramatically shift your party's power curve, making them high-priority targets for any serious build.

S-Tier Drops

• Chromatic Greatsword: Exclusive to the Chromatic Greatsword Cultist, this weapon scales with Vitality and Agility for Sireso, making it ideal for hybrid tank-damage builds.
• Reacher: A top-tier choice for Tireso that synergizes well with high-crit loadouts.
• Ballaro: Perfect for Monoco thanks to dual scaling with Agility and Defense, turning the character into a durable front-liner.

A-Tier Drops

• Angerim: High raw damage output, best used on burst-damage characters.
• Gobluson: Balanced stats make it a reliable pick for mid-game progression.
• Bourgelon: Excels in sustained fights due to its built-in lifesteal mechanic.

B-Tier Drops

• Lusteson: Situational, but shines in elemental-heavy encounters.
• Benisim: Offers utility over raw power, useful for support builds.
• Facesum: Niche pick for crowd-control focused teams.
• Jarum: Decent early-game upgrade, quickly outclassed by S-tier options.

Best Pictos Tier List

Pictos dropped by Chromatic Nevrons provide powerful passive buffs that can define entire playstyles. Prioritize these based on your party composition and elemental focus.

S-Tier Utility

• Anti-Stun: Grants complete immunity to stun effects, invaluable against elite packs.
• Anti-Blight: Prevents Blight status, a game-changer in poison-heavy zones.

A-Tier Synergy

• Energising Burn: Core for Burn-based Maelle builds, amplifying damage over time.
• First Offensive: Boosts opening burst damage, ideal for speedrun strategies.
• Longer Burn: Extends Burn duration, stacking well with other fire synergies.

B-Tier Situational

• Powerful Revive: Increases revive potency, useful for glass-cannon teams.
• Shielding Death: Grants a temporary shield on death, offering a safety net for risky plays.

Essential Upgrade Materials

Beyond weapons and Pictos, Chromatic Nevrons drop rare materials crucial for endgame progression. These resources are the bottleneck for reaching maximum power.

Level 32 Breakthrough

• Grandiose Chroma Catalyst: Dropped frequently by most Chromatic Nevrons, this is the key material needed to push any weapon to Level 32.

Level 33 Capstone

• Perfect Chroma Catalyst: A single guaranteed drop from the Chromatic Création Nevron, required to achieve the ultimate Level 33 upgrade for one weapon.

Respec & Mastery

• Colour of Lumina: Common drop from Chromatic Goblu, used to respec Skill Points and gain additional Lumina Points for late-game flexibility.

Community Tips and Reddit Strategies

Finding Hidden Chromatic Nevrons

The r/expedition33 community has rallied around a single thread to track every Chromatic Nevron—elite optional bosses scattered across the Continent. Nearly 50 variants exist, and most are tucked away far from the critical path.

• Environmental Secrets: Check behind illusory walls and breakable surfaces. If a wall looks odd, hit it.
• Interactive Map: Cross-reference your progress with the community-maintained map to avoid missing the sneakiest spawns.
• Missing Entries: Six enemy types—Contorsionniste, Dualliste, Gargant, Pèlerin, Stalact, and Volester—currently have no confirmed Chromatic forms, so don't waste time hunting them.

Optimal Party Compositions

Reddit consensus is clear: synergy beats raw stats. The "perfect" party shifts with every act, boss, and objective.

Early-Game Staples

• Tank: Draws aggro and soaks damage.
• Healer/Support: Keeps everyone upright.
• Damage Dealer: Focuses on elemental weaknesses.

Mid-to-Late Game Specializations

• Stance-Breaking Teams: Rapidly crack enemy defenses for devastating follow-ups.
• Buff Stacking/Debuff Teams: Layer vulnerability effects, then detonate them for spike damage.

Remember: the "best" squad is situational. Build for area clears, single-target burst, or specific mechanics—never one-size-fits-all.

Cheese Strategies and One-Shot Builds

When brute force fails, the community turns to cheese builds—high-risk, high-reward setups that can erase even the toughest bosses in a single turn.

Top-Tier One-Shotters

• Maelle: Equip Medalum (Level 33) and stack every Lumina that boosts damage or crit damage.
• Verso: Combine free-aim builds with Mark and Burn debuffs for rapid takedowns on bosses like Osquio.

Execution Flow

1. Support Units apply Mark and Burn.
2. Stance Break the target.
3. Primary Damage Dealer unleashes the perfected rotation.

Some players call these tactics a "necessary evil" for New Game+ and post-game content—because when conventional strategies hit a wall, cheese cuts straight through.

Chromatic Nevron Checklist

Track your progress against all Chromatic Nevrons: optional end-game bosses that drop some of the best gear in Clair Obscur: Expedition 33. Use the table below to mark each encounter as you complete it.

Printable Checklist

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Quick Reference Notes

• Total Chromatic Nevrons: ~50 scattered across all three Acts
• Key Rewards: Grandiose Chroma Catalysts, Colours of Lumina, top-tier weapons and Pictos
• Required Abilities: Some need Esquie's flight or underwater diving
• Trophy: Defeating them all unlocks the "Aiding the Enemy" achievement

Hunting Chromatic Nevrons is the definitive path to power in Expedition 33, rewarding you with top-tier weapons, essential Pictos, and rare upgrade materials. With nearly 50 of these formidable bosses to track, use this guide to systematically conquer each encounter and complete your collection. Now, gear up and prove your mastery across the Continent.